Sindarin Phonetic Development (Part 72)

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S. short vowels vanished before morpheme boundaries; [-C{ĭĕăŏ}] > [-Cø]

Just as short final vowels vanished in Sindarin and Noldorin at the end of words, they also vanished in the middle of words at morpheme boundaries. As suggested by David Salo (GS/§4.115), this vowel loss at morpheme boundaries was later than the loss at the end of words. There are a couple of examples that support this ordering of sound changes:

Sindarin Phonetic Development (Part 70)

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S. [ɣ] became [i] between sonants and vowels; [{lrð}ɣV] > [{lrð}iV]

Where [ɣ] appears between a sonant (voiced consonants like [l], [r] and [ð]) and a vowel, it changed into the vowel [i], as noted by David Salo (GS/§4.152). All the examples we have of this phonetic development are from Noldorin, but we have no reason to believe Sindarin was any different:

Sindarin Phonetic Development (Part 65)

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S. [ę̄] became [ai]; [ę̄] > [ai]

The Primitive Elvish long vowel [ę̄], a slacker and lower long [ē], developed into [ai] in Sindarin (PE18/96). A similar sound change occurred in Noldorin, except at that conceptual stage Tolkien represented the sound as [ǣ], with a slightly different pronunciation. Tolkien described this sound change in several places, with somewhat different details each time:

Sindarin Phonetic Development (Part 64)

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S. [ǭ] became [au]; [ǭ] > [au]

The Primitive Elvish long vowel [ǭ], a slacker and lower long [ō], developed into [au] in Sindarin (PE18/96). A similar sound change occurred in Noldorin, though the exact details of the phonetic development are different. Tolkien described this sound change in several places, with somewhat different details each time: